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(Coleoptera, Diptera, Hymenoptera and Lepidoptera)
by
Brian Pitkin, Willem Ellis, Colin Plant and Rob Edmunds
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MELICA.
Slough-grasses. [Poaceae]
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Four
species of Melica are recorded in Britain. These include
the native Mountain Melick (M. nutans) and the Wood Melick
(M. uniflora).
Thirteen British miners are recorded on Melica.
Nearly 100 British miners or possible miners are recorded on grasses in Britain.
A key to the European miners recorded on Melica is provided in Bladmineerders van Europa.
It is recommended that adults of all miners on grasses be reared to be certain of their identity. |
Key for the identification of the known mines of British
insects (Diptera and non-Diptera) recorded on Melica
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1# > Leaf-miner: Details unknown. Probably the larva mines for a few days only, and afterwards lives as a stem borer. |
On Poaceae in Britain, On Bromus, Elytrigia, Hordeum, Melica, Secale and Triticum elswhere. Widespread in Britain and continetal Europe. Also
recorded in the Republic of Ireland and Northern Ireland
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Ochsenheimeria
urella Fischer von Röslerstamm,
1842 [Lepidoptera:
Ypsolophidae]. |
1a > Leaf-miner: Tufted
hair-grass and blue moor-grass are the main foodplants, the larvae
forming gallery mines.
Gradually
widening corridor, running either upwards or down. All frass is
deposited in the earliest part of the mine. Often 2-3 larvae in
a mine; in grasses with broad leaves sometimes more than one mine
in a leaf. |
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On Deschampsia and Sesleria, but not yet on Melica,
in Britain and Carex, Brachypodium, Calamagrostis, Deschampsia,
Elymus, Festuca, Melica, Milium, Phleum, Poa and Sesleria elsewhere. Occurs in woodland habitats in England, Wales and locally
in Ireland. Also recorded in the Channel Is. and the Republic
of Ireland. Widespread in continental Europe.
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Elachista
adscitella Stainton, 1851 [Lepidoptera: Elachistidae]. |
1b > Leaf-miner: Corridor, 15 cm in the end, descending from the leaf tip. The mine
is whtish and shallow at first, then becomes deeper, yellowish white,
and more transparant. Pupation external; pupa, not in a cocoon,
attached to the leaf. |
On Carex, Eleocharis and Eriophorum, but not yet on Melica, in Britain and Carex, Eleocharis,
Eriophorum, Scirpus, Calamagrostis, Deschampsia, Melica and Poa elsewhere. Northern Britain & Ireland. Also recorded
in the Republic of Ireland. Widespread in continental Europe.
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Elachista
albidella Nylander, 1848 [Lepidoptera: Elachistidae]. |
1c > Leaf-miner: Corridor
widening while descending from the tip of the leaf. The mine is
unusual because the sides are very irregularly scalloped out. Moreover,
the mine is not evenly transparent, but rather yellowish green and
motly, because the larva leaves patches of parenchyma uneaten, and
does not feed full depth. Frass in a few irregular, interrupted
length lines. Often 2-3 larvae in a mine. The larvae hibernate in
the centre of the mine; after winter they leave their mine and pupate. |
On 'various grasses', but not yet on Melica, in
Britain plus Luzula, Agrostis, Arrhenatherum, Brachypodium,
Calamagorstis, Dactylis, Deschampsia, Elymus, Festuca, Glyceria,
Holcus, Melica, Milium and Poa elsewhere. Widespread
in Britain, Ireland and continental Europe.
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Elachista
apicipunctella Stainton, 1849 [Lepidoptera: Elachistidae]. |
1d > Leaf-miner: Like related species, the larva mines blades of grass, in this case
usually Cock's-foot, forming a thin whitish mine.
The
larva begins in autumn the making of a long, narrow, corridor with
a fine central line of grey frass. The corridor is straight or lightly
wavy, and descends into the leaf sheath, or even into the stem or
rootstock. The larva regularly leaves the mine to begin making a
new one. Pupation external. |
On Dactylis, but not yet on Melica, in Britain
and Bromus, Carex, Dactylis, Melica and Milium elsewhere.
Widespread in Britain, Ireland and continental Europe.
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Elachista
atricomella Stainton, 1849 [Lepidoptera: Elachistidae]. |
1e > Leaf-miner: Mine generally descending from the leaf tip. The mine may occupy
the space between leaf margin and midrib, but may also occupy the
entire width of the leaf. Most frass accumulated in the oldest part
of the mine. Pupation external; the pupa is attached to the leaf
without a cocoon. Larval head and prothoracic shield dark brown, ody yellowish white, with a pair of orange red latero-dorsal length
lines; ventrally another such line, medially. |
On Carex and Deschampsia, but not yet on Melica,
in Britain plus Melica and Sesleria elsewhere. Britain
and Northern Ireland. Also recorded in the Republic of Ireland.
Widespread in continental Europe.
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Elachista
cinereopunctella (Haworth, 1828) [Lepidoptera: Elachistidae]. |
1f > Leaf-miner: The larva mines downwards and forms an irregular mine with a silken
tube in the centre, which is mixed with frass.
Mine
transparent (therefore conspicuous), generally descending from the
leaf tip. Over the entire length of the mine stretches a central
silken tube, in which the larva can retreat and can move quickly
up or down. The tube also contains the frass. The larva feeds laterally
from the tube, which makes the sides of the mine very irregular. |
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On Brachypodium, Dactylis and Holcus, but not yet on Melica, in Britain and Brachypodium, Dactylis and Melica elsewhere. Distributed in southern England and
Wales and also occurring in parts of northern England and Ireland.
Widespread in continental Europe.
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Elachista
gangabella Zeller, 1850 [Lepidoptera: Elachistidae]. |
1g > Leaf-miner: Long,
narrow, white corridor, descending from close to the leaf tip to
the leaf base or even stem. Frass in an inconspicuous grey line.
From the stem the larva may enter a new leaf (Steuer, 1987a; Bland,
1996a). |
On Brachypodium and Dactylis, but not yet on Melica, in Britain and Bromopsis, Dactylis, Deschampsia,
Festuca, Lolium, Melica, Milium and Poa elsewhere.
Widespread in Britain, Ireland and continental Europe.
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Elachista
luticomella Zeller, 1839 [Lepidoptera: Coleophoridae]. |
1h > Leaf-miner: Initially a narrow brownish mine with blackish frass at its base,
then moves to another leaf, forming a broader mine. Both mines can
pucker the blade. |
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On Brachypodium and Bromopsis in Britain and numerous grasses and sedges in continetal Europe.Southern half of England. Also recorded in
the Republic of Ireland. Widespread in continental Europe.
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Elachista
obliquella Stainton, 1858 [Lepidoptera: Elachistidae]. |
1i > Leaf-miner: The
larva creates a long narrow mine in the blade.
Mine
fairly translucent. It starts as a narrow corridor, running towards
the leaf tip, but this almost invariably is overrun be a later,
broad, descending corridor. The centre of the mine contains a vertical
tube made of loose silk, in which the larva can quickly move up
or down, and in which also most frass gets trapped. The sides of
the mine are smooth (contrary to E.
gangabella). Mines are difficult to find, because the leaves
are wilting when larvae are present. |
On Molinia, but not yet on Melica, in Britain
and Carex, Arrhenatherum, Brachypodium, Calamagrostis, Melica,
Molinia and Poa elsewhere.
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Elachista
subalbidella Schläger, 1847 [Lepidoptera: Elachistidae]. |
1j > Leaf-miner: Broad
elongated mine; the form is dependent of the leaf form of the host
plant. Frass green. Usually a number of larvae together in a mine.
Pupation in the mine. |
On Bromopsis, Dactylis, Elymus and Phalaris, but not yet on Melica, in Britain
and additional grasses, including Elytrigia, elsewhere. Widespread in Britain. Also
recorded in the Republic of Ireland. Widespread and common in continental Europe. Also recorded in Japan, U.S.A. and Canada.
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Cerodontha
incisa (Meigen, 1830) [Diptera: Agromyzidae]. |
1k > Leaf-miner: Normally
several larvae feed together. Pupation in the mine (Spencer,
1976: 198).
Broad
elongated blotch. Frass greenish. Larvae generally communal. Pupation
within the mine. The black puaria are individially anchored within
the mine with a silken thread attached at their rear end. Distinguishable
from C. incisa only
by means of the larva. |
On Arrhenatherum, Brachypodium, Bromus, Bromopsis, Calamagrostis,
Dactylis, Deschampsia, Elymus, Festuca, Holcus, Lolium, Molinia and Phalaris, but not yet on Melica, in Britain and
additional grasses elsewhere. Widespread and common in Britain.
Also recorded in the Republic of Ireland. Widespread and frequently
common in much of Europe. Also recorded in Alaska and Canada.
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Cerodontha
pygmaea (Meigen, 1830) [Diptera: Agromyzidae]. |
1l > Leaf-miner: Narrow whitish mine, with frass in distinct black lumps. Pupation
internal (Spencer, 1976:
422).
Whitish,
upper-surface, descending corridor, about halfway up the blade.
Frass in distinct black grains that are lying further apart than
their diameter. Pupation in the mine. |
On Arrhenatherum, Holcus, Milium and Phleum, but not yet on Melica, in Britain and additional grasses elsewhere. Recorded in Scotland
and Widespread in continental Europe. Also recorded in Canada.
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Chromatomyia
fuscula (Zetterstedt, 1838) [Diptera: Agromyzidae]. |
1m > Leaf-miner: Long, narrow, whitish mine. Pupation internal (Spencer,
1976: 453); anterior spiracles projecting through the epidermis.
Whitish,
upper-surface, rather narrow corridor with comparatively large frass
grains that are laying further apart than their diameter. Pupation
within the mine. The anterior spiracles of the orange-brown puparium
penetrate the epidermis. |
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Chromatomyia nigra larva, lateral
Image: © Willem Ellis (Bladmineerders van Europa) |

Chromatomyia nigra pupa, lateral
Image: © Willem Ellis (Bladmineerders van Europa) |
On
numerous genera of grasses, but not yet on Melica, in Britain. Widespread
and common throughout British Isles and much of Europe. Also recorded
in Canada, western U.S.A. and Japan.
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Chromatomyia
nigra (Meigen, 1830) [Diptera: Agromyzidae]. |
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